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<article article-type="research-article" dtd-version="1.3" xmlns:mml="http://www.w3.org/1998/Math/MathML" xmlns:xlink="http://www.w3.org/1999/xlink" xmlns:xsi="http://www.w3.org/2001/XMLSchema-instance" xml:lang="en"><front><journal-meta><journal-id journal-id-type="publisher-id">gesj</journal-id><journal-title-group><journal-title xml:lang="en">GEOGRAPHY, ENVIRONMENT, SUSTAINABILITY</journal-title><trans-title-group xml:lang="ru"><trans-title>GEOGRAPHY, ENVIRONMENT, SUSTAINABILITY</trans-title></trans-title-group></journal-title-group><issn pub-type="ppub">2071-9388</issn><issn pub-type="epub">2542-1565</issn><publisher><publisher-name>Russian Geographical Society</publisher-name></publisher></journal-meta><article-meta><article-id pub-id-type="doi">10.24057/2071-9388-2018-11-3-21-38</article-id><article-id custom-type="elpub" pub-id-type="custom">gesj-473</article-id><article-categories><subj-group subj-group-type="heading"><subject>Research Article</subject></subj-group><subj-group subj-group-type="section-heading" xml:lang="en"><subject>GEOGRAPHY</subject></subj-group><subj-group subj-group-type="section-heading" xml:lang="ru"><subject>География</subject></subj-group></article-categories><title-group><article-title>MIDDLE PLEISTOCENE SMALL MAMMAL fAUNAS Of EUROPE: EVOLUTION, BIOSTRATIGRAPHY, CORRELATIONS</article-title><trans-title-group xml:lang="ru"><trans-title></trans-title></trans-title-group></title-group><contrib-group><contrib contrib-type="author" corresp="yes"><name-alternatives><name name-style="western" xml:lang="en"><surname>Markova</surname><given-names>A. K.</given-names></name></name-alternatives><bio xml:lang="en"><p>Doctor of Geographical Sciences.</p><p>Moscow.</p></bio><email xlink:type="simple">amarkova@list.ru</email><xref ref-type="aff" rid="aff-1"/></contrib><contrib contrib-type="author" corresp="yes"><name-alternatives><name name-style="western" xml:lang="en"><surname>Puzachenko</surname><given-names>A. Yu.</given-names></name></name-alternatives><bio xml:lang="en"><p>Doctor of biological Sciences.</p><p>Moscow.</p></bio><xref ref-type="aff" rid="aff-1"/></contrib></contrib-group><aff xml:lang="en" id="aff-1"><institution>Institute of Geography of the Russian Academy of Sciences.</institution><country>Russian Federation</country></aff><pub-date pub-type="collection"><year>2018</year></pub-date><pub-date pub-type="epub"><day>29</day><month>09</month><year>2018</year></pub-date><volume>11</volume><issue>3</issue><fpage>21</fpage><lpage>38</lpage><permissions><copyright-statement>Copyright &amp;#x00A9; Markova A.K., Puzachenko A.Y., 2018</copyright-statement><copyright-year>2018</copyright-year><copyright-holder xml:lang="ru">Markova A.K., Puzachenko A.Y.</copyright-holder><copyright-holder xml:lang="en">Markova A.K., Puzachenko A.Y.</copyright-holder><license xml:lang="ru" license-type="creative-commons-attribution" xlink:href="https://creativecommons.org/licenses/by/4.0/" xlink:type="simple"><license-p>Данная работа распространяется под лицензией Creative Commons Attribution 4.0.</license-p></license><license xml:lang="en" license-type="creative-commons-attribution" xlink:href="https://creativecommons.org/licenses/by/4.0/" xlink:type="simple"><license-p>This work is licensed under a Creative Commons Attribution 4.0 License.</license-p></license></permissions><self-uri xlink:href="https://ges.rgo.ru/jour/article/view/473">https://ges.rgo.ru/jour/article/view/473</self-uri><abstract><p>The paper is concerned with the small mammal fauna evolution in Europe in the Middle Pleistocene. The information on the faunas of the end of the Early Pleistocene has been also taken into consideration. The data available made possible identifying several stages in the small mammal evolution. Not all intervals within the Middle Pleistocene are provided with sufficient information for recognizing individual stages; that is particularly true for the cold periods of the Middle Pleistocene – the Donian and the Okian glaciations (=Elsterian, =Anglian). Based on the studies of small mammal localities, the biostratigraphic scheme has been developed, the principal phylogenetic lineages of Arvicolinae were traced, and maps of the Middle Pleistocene small mammal localities have been compiled</p></abstract><kwd-group xml:lang="en"><kwd>Middle Pleistocene</kwd><kwd>small mammals</kwd><kwd>Europe</kwd><kwd>biostratigraphy</kwd><kwd>evolution</kwd><kwd>correlations</kwd></kwd-group></article-meta></front><back><ref-list><title>References</title><ref id="cit1"><label>1</label><citation-alternatives><mixed-citation xml:lang="ru">Agadjanian A.K. (2009). Small mammals of the Pliocene – Pleistocene of the Russian Plain. Moscow: Nauka Press (in Russian).</mixed-citation><mixed-citation xml:lang="en">Agadjanian A.K. (2009). Small mammals of the Pliocene – Pleistocene of the Russian Plain. Moscow: Nauka Press (in Russian).</mixed-citation></citation-alternatives></ref><ref id="cit2"><label>2</label><citation-alternatives><mixed-citation xml:lang="ru">Agusti J. (1992).The Allophaiomys complex in Southern Europe, Geobios, 25 (1), pp. 133-144 Alexandrova L.P. (1976).</mixed-citation><mixed-citation xml:lang="en">Agusti J. (1992).The Allophaiomys complex in Southern Europe, Geobios, 25 (1), pp. 133-144 Alexandrova L.P. (1976).</mixed-citation></citation-alternatives></ref><ref id="cit3"><label>3</label><citation-alternatives><mixed-citation xml:lang="ru">Anthropogene rodents of the European part of USSR. Moscow: Nauka Press (in Russian).</mixed-citation><mixed-citation xml:lang="en">Anthropogene rodents of the European part of USSR. Moscow: Nauka Press (in Russian).</mixed-citation></citation-alternatives></ref><ref id="cit4"><label>4</label><citation-alternatives><mixed-citation xml:lang="ru">Bates M.R., Wenban-Smith F.F., Bello S.M., Bridgland D.R., Buck L.T., Collins M.J., Keen, J., Leary D.H., Parfitt S.A., Penkman K., Rhodes E., Ryssaer C. and Whittaker J.E. (2014). Late persistence of the Acheulian in southern Britain in an MIS 8 interstadial: evidence from Harnham, Wiltshire. Quaternary Science Reviews, 101, pp. 159-176.</mixed-citation><mixed-citation xml:lang="en">Bates M.R., Wenban-Smith F.F., Bello S.M., Bridgland D.R., Buck L.T., Collins M.J., Keen, J., Leary D.H., Parfitt S.A., Penkman K., Rhodes E., Ryssaer C. and Whittaker J.E. (2014). Late persistence of the Acheulian in southern Britain in an MIS 8 interstadial: evidence from Harnham, Wiltshire. Quaternary Science Reviews, 101, pp. 159-176.</mixed-citation></citation-alternatives></ref><ref id="cit5"><label>5</label><citation-alternatives><mixed-citation xml:lang="ru">Blackwell B. and Schwarcz H.P. (1986). U-Series analyses of the Lower Travertine at Ehringsdorf, DDR. Quaternary Research, 25 (2), pp. 215-222.</mixed-citation><mixed-citation xml:lang="en">Blackwell B. and Schwarcz H.P. (1986). U-Series analyses of the Lower Travertine at Ehringsdorf, DDR. Quaternary Research, 25 (2), pp. 215-222.</mixed-citation></citation-alternatives></ref><ref id="cit6"><label>6</label><citation-alternatives><mixed-citation xml:lang="ru">Chaline J. (1972). Les Rongeurs du Pleistocéne moyen et supérieur de la France. Paris: Cahiers de Paléontologie, CNRF.</mixed-citation><mixed-citation xml:lang="en">Chaline J. (1972). Les Rongeurs du Pleistocéne moyen et supérieur de la France. Paris: Cahiers de Paléontologie, CNRF.</mixed-citation></citation-alternatives></ref><ref id="cit7"><label>7</label><citation-alternatives><mixed-citation xml:lang="ru">Cuenca-Bescós G., Rofes J., López-García J.-M., Blain H.-A., Rabal-Garcés R., Sauqué V., Arsuaga J.L., Bermúdez de Castro J. M. and Carbonell, E. (2013). The small mammals of Sima del Elefante (Atapuerca, Spain) and the first entrance of Homo in Western Europe. Quaternary International. 295, pp. 28–35.</mixed-citation><mixed-citation xml:lang="en">Cuenca-Bescós G., Rofes J., López-García J.-M., Blain H.-A., Rabal-Garcés R., Sauqué V., Arsuaga J.L., Bermúdez de Castro J. M. and Carbonell, E. (2013). The small mammals of Sima del Elefante (Atapuerca, Spain) and the first entrance of Homo in Western Europe. Quaternary International. 295, pp. 28–35.</mixed-citation></citation-alternatives></ref><ref id="cit8"><label>8</label><citation-alternatives><mixed-citation xml:lang="ru">Cyrek K., Socha P., Stefaniak K., Madeyska T., Miroslaw-Grabowska J., Sudoł M. and Czyźewski Ł. (2010). Palaeolithic of Biśnic Cave (Southern Poland) within the environmental Background. Quaternary International. 220, pp. 5–30.</mixed-citation><mixed-citation xml:lang="en">Cyrek K., Socha P., Stefaniak K., Madeyska T., Miroslaw-Grabowska J., Sudoł M. and Czyźewski Ł. (2010). Palaeolithic of Biśnic Cave (Southern Poland) within the environmental Background. Quaternary International. 220, pp. 5–30.</mixed-citation></citation-alternatives></ref><ref id="cit9"><label>9</label><citation-alternatives><mixed-citation xml:lang="ru">Gibbard P.L., Cohen K.M., Boreham S. and Moscariello A. (2004). Global chronostratigraphical correlation table for the last 2.7 million years. Subcommission on Quaternary Stratigraphy, International Commission on Stratigraphy: Cambridge.</mixed-citation><mixed-citation xml:lang="en">Gibbard P.L., Cohen K.M., Boreham S. and Moscariello A. (2004). Global chronostratigraphical correlation table for the last 2.7 million years. Subcommission on Quaternary Stratigraphy, International Commission on Stratigraphy: Cambridge.</mixed-citation></citation-alternatives></ref><ref id="cit10"><label>10</label><citation-alternatives><mixed-citation xml:lang="ru">Gromov V.I. (1948). Palaeontological and archaeological foundation of the stratigraphy of the Quaternary continental deposits in the territory of the USSR. Moscow: the USSR Academy of Sciences (in Russian).</mixed-citation><mixed-citation xml:lang="en">Gromov V.I. (1948). Palaeontological and archaeological foundation of the stratigraphy of the Quaternary continental deposits in the territory of the USSR. Moscow: the USSR Academy of Sciences (in Russian).</mixed-citation></citation-alternatives></ref><ref id="cit11"><label>11</label><citation-alternatives><mixed-citation xml:lang="ru">Grün R. and Schwartz H.P. (2000). Revised open system U-series/ESR age calculations for teeth from Stratum C at the Hoxnian interglacial type locality, England. Quaternary Science Reviews, 19, pp.1151-1154.</mixed-citation><mixed-citation xml:lang="en">Grün R. and Schwartz H.P. (2000). Revised open system U-series/ESR age calculations for teeth from Stratum C at the Hoxnian interglacial type locality, England. Quaternary Science Reviews, 19, pp.1151-1154.</mixed-citation></citation-alternatives></ref><ref id="cit12"><label>12</label><citation-alternatives><mixed-citation xml:lang="ru">Heinrich W.-D. (1978). Zur biometrischen Erfassung eines Evolutionstrends bei Arvicola (Rodentia, Mammalia) aus dem Pleistozän Thüringens. Berlin: Säugetierkdl. Inform., 2, pp. 3-21.</mixed-citation><mixed-citation xml:lang="en">Heinrich W.-D. (1978). Zur biometrischen Erfassung eines Evolutionstrends bei Arvicola (Rodentia, Mammalia) aus dem Pleistozän Thüringens. Berlin: Säugetierkdl. Inform., 2, pp. 3-21.</mixed-citation></citation-alternatives></ref><ref id="cit13"><label>13</label><citation-alternatives><mixed-citation xml:lang="ru">Heinrich W.-D. (1990). Some aspects of evolution and biostratigraphy of Arvicola (Mammalia, Rodentia) in the central European Pleistocene. Prague: Int. Symp. Evol. Phyl. Biostr. Arvicolids., pp. 165-180.</mixed-citation><mixed-citation xml:lang="en">Heinrich W.-D. (1990). Some aspects of evolution and biostratigraphy of Arvicola (Mammalia, Rodentia) in the central European Pleistocene. Prague: Int. Symp. Evol. Phyl. Biostr. Arvicolids., pp. 165-180.</mixed-citation></citation-alternatives></ref><ref id="cit14"><label>14</label><citation-alternatives><mixed-citation xml:lang="ru">Fejfar O. and Horaček I. (1990). Review of fossil arvicolids (Mammalia, Rodentia) of the Pliocene and Quaternary in Czechoslovakia. Int. Symp. Evol. Phyl. Biostr. Arvicolids. Prague: Geological Survey, pp. 125–132.</mixed-citation><mixed-citation xml:lang="en">Fejfar O. and Horaček I. (1990). Review of fossil arvicolids (Mammalia, Rodentia) of the Pliocene and Quaternary in Czechoslovakia. Int. Symp. Evol. Phyl. Biostr. Arvicolids. Prague: Geological Survey, pp. 125–132.</mixed-citation></citation-alternatives></ref><ref id="cit15"><label>15</label><citation-alternatives><mixed-citation xml:lang="ru">Iosifova Yu.I., Agadjanian A.K., Pisareva V.V., and Semenov V.V. (2006). The Upper Don basin is the stratotype of the Middle Pleistocene of Russian Plain. Palynological Reconstructions: Stratigraphy, Geoecology. S.-Petersburg: Nedra Press, pp. 41–121 (in Russian) .</mixed-citation><mixed-citation xml:lang="en">Iosifova Yu.I., Agadjanian A.K., Pisareva V.V., and Semenov V.V. (2006). The Upper Don basin is the stratotype of the Middle Pleistocene of Russian Plain. Palynological Reconstructions: Stratigraphy, Geoecology. S.-Petersburg: Nedra Press, pp. 41–121 (in Russian) .</mixed-citation></citation-alternatives></ref><ref id="cit16"><label>16</label><citation-alternatives><mixed-citation xml:lang="ru">Krasnenkov R.V. and Kazantceva, N.E. (1993). The discovery of the early Dnieper alluvium in the Upper Don terraces, Bull. of regional commission of Central and Southern Russian Platform, 2, pp.153-161 (in Russian).</mixed-citation><mixed-citation xml:lang="en">Krasnenkov R.V. and Kazantceva, N.E. (1993). The discovery of the early Dnieper alluvium in the Upper Don terraces, Bull. of regional commission of Central and Southern Russian Platform, 2, pp.153-161 (in Russian).</mixed-citation></citation-alternatives></ref><ref id="cit17"><label>17</label><citation-alternatives><mixed-citation xml:lang="ru">Krajcarz M.T. (2012). Small fossil wolverine Gulo from Middle Pleistocene of Poland. Acta zoologica cracoviensia, 55(1), pp. 79-87.</mixed-citation><mixed-citation xml:lang="en">Krajcarz M.T. (2012). Small fossil wolverine Gulo from Middle Pleistocene of Poland. Acta zoologica cracoviensia, 55(1), pp. 79-87.</mixed-citation></citation-alternatives></ref><ref id="cit18"><label>18</label><citation-alternatives><mixed-citation xml:lang="ru">Krokhmal A.I. and Rekovets L.I. (2010). Pleistocene small mammal localities of Ukraine and adjacent territories. Kiev: LAT and K Press (In Russian)</mixed-citation><mixed-citation xml:lang="en">Krokhmal A.I. and Rekovets L.I. (2010). Pleistocene small mammal localities of Ukraine and adjacent territories. Kiev: LAT and K Press (In Russian)</mixed-citation></citation-alternatives></ref><ref id="cit19"><label>19</label><citation-alternatives><mixed-citation xml:lang="ru">Lisiecki L.E. and Raymo M.E. (2005). A Pliocene-Pleistocene stack of 57 globally distributed benthic δ18O records. Paleoceanography, 20 (1), PA1003. doi:10.1029/2004PA001071.</mixed-citation><mixed-citation xml:lang="en">Lisiecki L.E. and Raymo M.E. (2005). A Pliocene-Pleistocene stack of 57 globally distributed benthic δ18O records. Paleoceanography, 20 (1), PA1003. doi:10.1029/2004PA001071.</mixed-citation></citation-alternatives></ref><ref id="cit20"><label>20</label><citation-alternatives><mixed-citation xml:lang="ru">Markova A.K. (1975). Pleistocene rodents of Dnieper basin and its significance for the palaeogeography and stratigraphy. Aftoref. disser. na soisk. uch. stepeni kand. geogr. nauk. Moscow: Institute of Geogr. Ac. Nauk USSR Press (in Russian).</mixed-citation><mixed-citation xml:lang="en">Markova A.K. (1975). Pleistocene rodents of Dnieper basin and its significance for the palaeogeography and stratigraphy. Aftoref. disser. na soisk. uch. stepeni kand. geogr. nauk. Moscow: Institute of Geogr. Ac. Nauk USSR Press (in Russian).</mixed-citation></citation-alternatives></ref><ref id="cit21"><label>21</label><citation-alternatives><mixed-citation xml:lang="ru">Markova A.K. (1981). Morphological changes of water voles teeth during the Pleistocene, In: A.A.Velichko and S.M. Shik, eds., The questions in Pleistocene palaeogeography of glacial and periglacial regions. Moscow: Nauka Press, pp. 91-110 (in Russian).</mixed-citation><mixed-citation xml:lang="en">Markova A.K. (1981). Morphological changes of water voles teeth during the Pleistocene, In: A.A.Velichko and S.M. Shik, eds., The questions in Pleistocene palaeogeography of glacial and periglacial regions. Moscow: Nauka Press, pp. 91-110 (in Russian).</mixed-citation></citation-alternatives></ref><ref id="cit22"><label>22</label><citation-alternatives><mixed-citation xml:lang="ru">Markova A.K. (1982). Pleistocene rodents of the Russian Plain. Moscow: Nauka Press (in Russian).</mixed-citation><mixed-citation xml:lang="en">Markova A.K. (1982). Pleistocene rodents of the Russian Plain. Moscow: Nauka Press (in Russian).</mixed-citation></citation-alternatives></ref><ref id="cit23"><label>23</label><citation-alternatives><mixed-citation xml:lang="ru">Markova A.K. (1990). The sequence of Early Pleistocene small-mammal faunas from the South Russian Plain. Quartärpaläontologie, 8, pp. 131-151.</mixed-citation><mixed-citation xml:lang="en">Markova A.K. (1990). The sequence of Early Pleistocene small-mammal faunas from the South Russian Plain. Quartärpaläontologie, 8, pp. 131-151.</mixed-citation></citation-alternatives></ref><ref id="cit24"><label>24</label><citation-alternatives><mixed-citation xml:lang="ru">Markova A.K. (1992). Pleistocene microtheriofauna of Eastern Europe. In: A.A. Velichko and S.M. Shik, eds., Stratigraphy and Paleogeography of Quaternary of Eastern Europe, Moscow: Inst. of Geography RAS, pp. 50-94 (in Russian).</mixed-citation><mixed-citation xml:lang="en">Markova A.K. (1992). Pleistocene microtheriofauna of Eastern Europe. In: A.A. Velichko and S.M. Shik, eds., Stratigraphy and Paleogeography of Quaternary of Eastern Europe, Moscow: Inst. of Geography RAS, pp. 50-94 (in Russian).</mixed-citation></citation-alternatives></ref><ref id="cit25"><label>25</label><citation-alternatives><mixed-citation xml:lang="ru">Markova A.K. (1998). Pleistocene rodents of the Central and Southern Russian Plain. In: J.J. Saunders, B.W. Styles and G. F. Baryshnikov, eds., Quaternary Paleozoology in the Northern Hemisphere. Illinois State Museum Scientific papers, XXVI, pp. 119-143.</mixed-citation><mixed-citation xml:lang="en">Markova A.K. (1998). Pleistocene rodents of the Central and Southern Russian Plain. In: J.J. Saunders, B.W. Styles and G. F. Baryshnikov, eds., Quaternary Paleozoology in the Northern Hemisphere. Illinois State Museum Scientific papers, XXVI, pp. 119-143.</mixed-citation></citation-alternatives></ref><ref id="cit26"><label>26</label><citation-alternatives><mixed-citation xml:lang="ru">Markova A.K. (2005). Eastern European rodent (Rodentia, Mammalia) faunas from the EarlyMiddle Pleistocene transition. Quaternary International. 131, pp. 71-77.</mixed-citation><mixed-citation xml:lang="en">Markova A.K. (2005). Eastern European rodent (Rodentia, Mammalia) faunas from the EarlyMiddle Pleistocene transition. Quaternary International. 131, pp. 71-77.</mixed-citation></citation-alternatives></ref><ref id="cit27"><label>27</label><citation-alternatives><mixed-citation xml:lang="ru">Markova A.K. (2006). Likhvin Interglacial small mammal faunas of Eastern Europe. Quaternary International. 149 (1), pp. 67-79.</mixed-citation><mixed-citation xml:lang="en">Markova A.K. (2006). Likhvin Interglacial small mammal faunas of Eastern Europe. Quaternary International. 149 (1), pp. 67-79.</mixed-citation></citation-alternatives></ref><ref id="cit28"><label>28</label><citation-alternatives><mixed-citation xml:lang="ru">Markova A.K. (2007). Pleistocene mammal faunas of Eastern Europe. Quaternary International. 160, pp. 100-111.</mixed-citation><mixed-citation xml:lang="en">Markova A.K. (2007). Pleistocene mammal faunas of Eastern Europe. Quaternary International. 160, pp. 100-111.</mixed-citation></citation-alternatives></ref><ref id="cit29"><label>29</label><citation-alternatives><mixed-citation xml:lang="ru">Markova A.K. (2014). European small mammal faunas during the end of the Early Pleistocene – the beginning of the Middle Pleistocene. Izvestia RAS, Seria geograph., 5, pp. 83-98 (In Russian)</mixed-citation><mixed-citation xml:lang="en">Markova A.K. (2014). European small mammal faunas during the end of the Early Pleistocene – the beginning of the Middle Pleistocene. Izvestia RAS, Seria geograph., 5, pp. 83-98 (In Russian)</mixed-citation></citation-alternatives></ref><ref id="cit30"><label>30</label><citation-alternatives><mixed-citation xml:lang="ru">Markova A. and van Kolfschoten T. (2012). Middle Pleistocene small mammal faunas of Eastern and Western Europe: chronology, correlation. Geography, environment, sustainability, 5(4), pp. 17-23. Available at: https://doi.org/10.24057/2071-9388-2012-5-4-17-23.</mixed-citation><mixed-citation xml:lang="en">Markova A. and van Kolfschoten T. (2012). Middle Pleistocene small mammal faunas of Eastern and Western Europe: chronology, correlation. Geography, environment, sustainability, 5(4), pp. 17-23. Available at: https://doi.org/10.24057/2071-9388-2012-5-4-17-23.</mixed-citation></citation-alternatives></ref><ref id="cit31"><label>31</label><citation-alternatives><mixed-citation xml:lang="ru">Markova A.K. and Puzachenko A.Yu. (2016). The European small mammal faunas related to the first half of the Middle Pleistocene. Quaternary International. 420, pp. 378-390.</mixed-citation><mixed-citation xml:lang="en">Markova A.K. and Puzachenko A.Yu. (2016). The European small mammal faunas related to the first half of the Middle Pleistocene. Quaternary International. 420, pp. 378-390.</mixed-citation></citation-alternatives></ref><ref id="cit32"><label>32</label><citation-alternatives><mixed-citation xml:lang="ru">Markova A.K. and Vislobokova I.A. (2016). Mammal faunas in Europe at the end of the Early – beginning of the Middle Pleistocene. Quaternary International. 420, pp. 363- 377.</mixed-citation><mixed-citation xml:lang="en">Markova A.K. and Vislobokova I.A. (2016). Mammal faunas in Europe at the end of the Early – beginning of the Middle Pleistocene. Quaternary International. 420, pp. 363- 377.</mixed-citation></citation-alternatives></ref><ref id="cit33"><label>33</label><citation-alternatives><mixed-citation xml:lang="ru">Masini F. and Sala B. (2011). Considerations on an integrated biochronological scale of Italian Quaternary continental mammals. Italian Journal of Quaternary Sciences, 24 (2), pp. 193-198.</mixed-citation><mixed-citation xml:lang="en">Masini F. and Sala B. (2011). Considerations on an integrated biochronological scale of Italian Quaternary continental mammals. Italian Journal of Quaternary Sciences, 24 (2), pp. 193-198.</mixed-citation></citation-alternatives></ref><ref id="cit34"><label>34</label><citation-alternatives><mixed-citation xml:lang="ru">Maul L.C. (2000). Die Fossilfundstelle Weimar-Ehringsdorf. Eine Überisicht. Geowiss. Mitt. Thüringen, 10, pp. 167-174.</mixed-citation><mixed-citation xml:lang="en">Maul L.C. (2000). Die Fossilfundstelle Weimar-Ehringsdorf. Eine Überisicht. Geowiss. Mitt. Thüringen, 10, pp. 167-174.</mixed-citation></citation-alternatives></ref><ref id="cit35"><label>35</label><citation-alternatives><mixed-citation xml:lang="ru">Maul L.C. (2001). Die Kleinsäugerreste (Insectivora, Lagomorpha, Rodentia) aus dem Unterpleistozän von Untermassfeld. In: R.-D. Kahlke, ed., Das Pleistozän von Untermassfeld bei Meiningen (Thüringen). Monographien des Römisch-Germanischen Zentralmuseums Mainz, 40 (3), pp. 783–887.</mixed-citation><mixed-citation xml:lang="en">Maul L.C. (2001). Die Kleinsäugerreste (Insectivora, Lagomorpha, Rodentia) aus dem Unterpleistozän von Untermassfeld. In: R.-D. Kahlke, ed., Das Pleistozän von Untermassfeld bei Meiningen (Thüringen). Monographien des Römisch-Germanischen Zentralmuseums Mainz, 40 (3), pp. 783–887.</mixed-citation></citation-alternatives></ref><ref id="cit36"><label>36</label><citation-alternatives><mixed-citation xml:lang="ru">Maul L.C., Heinrich W.-D., Parfitt S.A. and Paunescu, A.-C. (2007). Comments on the correlation between magnetostratigraphy and evolution of Microtus (Arvicolidae, Rodentia, Mammalia) during the Early and early Middle Pleistocene. Courier Forschungsinstitut Senckenberg, 259, pp. 243-263.</mixed-citation><mixed-citation xml:lang="en">Maul L.C., Heinrich W.-D., Parfitt S.A. and Paunescu, A.-C. (2007). Comments on the correlation between magnetostratigraphy and evolution of Microtus (Arvicolidae, Rodentia, Mammalia) during the Early and early Middle Pleistocene. Courier Forschungsinstitut Senckenberg, 259, pp. 243-263.</mixed-citation></citation-alternatives></ref><ref id="cit37"><label>37</label><citation-alternatives><mixed-citation xml:lang="ru">Maul L. and Markova A.K. (2007). Similarity and regional differences in Quaternary arvicolid evolution in Central and Eastern Europe. Quaternary International. 160 (1), pp. 81-99.</mixed-citation><mixed-citation xml:lang="en">Maul L. and Markova A.K. (2007). Similarity and regional differences in Quaternary arvicolid evolution in Central and Eastern Europe. Quaternary International. 160 (1), pp. 81-99.</mixed-citation></citation-alternatives></ref><ref id="cit38"><label>38</label><citation-alternatives><mixed-citation xml:lang="ru">Maul L. and Parfitt S.A. (2010). Micromammals from the 1995 Mammoth Excavation at West Runton, Norfolk, UK: Morphometric data, biostratigraphy and taxonomic reappraisal. Quaternary International. 228, pp. 91-115.</mixed-citation><mixed-citation xml:lang="en">Maul L. and Parfitt S.A. (2010). Micromammals from the 1995 Mammoth Excavation at West Runton, Norfolk, UK: Morphometric data, biostratigraphy and taxonomic reappraisal. Quaternary International. 228, pp. 91-115.</mixed-citation></citation-alternatives></ref><ref id="cit39"><label>39</label><citation-alternatives><mixed-citation xml:lang="ru">Maul L.C., Rekovets L., Heinrich W.-D., Keller T. and Storch G. (2000). Arvicola mosbachensis (Schmidtgen 1911) of Mosbach 2: a basic sample for the early evolution of the genus and a reference for further biostratigraphical studies. Senckenbergiana lethaea, 80 (1), pp. 129-147.</mixed-citation><mixed-citation xml:lang="en">Maul L.C., Rekovets L., Heinrich W.-D., Keller T. and Storch G. (2000). Arvicola mosbachensis (Schmidtgen 1911) of Mosbach 2: a basic sample for the early evolution of the genus and a reference for further biostratigraphical studies. Senckenbergiana lethaea, 80 (1), pp. 129-147.</mixed-citation></citation-alternatives></ref><ref id="cit40"><label>40</label><citation-alternatives><mixed-citation xml:lang="ru">Mikhailesku C.D. and Markova A.K. (1992). The paleogeographical stages of fauna development on the south of Moldova in Anthropogene. Kishinev: Shtiintsa Press. (In Russian).</mixed-citation><mixed-citation xml:lang="en">Mikhailesku C.D. and Markova A.K. (1992). The paleogeographical stages of fauna development on the south of Moldova in Anthropogene. Kishinev: Shtiintsa Press. (In Russian).</mixed-citation></citation-alternatives></ref><ref id="cit41"><label>41</label><citation-alternatives><mixed-citation xml:lang="ru">Motuzko A.N. (1985). Anthropogene rodents of Belorussia and adjacent territories. Minsk: Nauka i Tekhnika Press, pp.173-186 (In Russian).</mixed-citation><mixed-citation xml:lang="en">Motuzko A.N. (1985). Anthropogene rodents of Belorussia and adjacent territories. Minsk: Nauka i Tekhnika Press, pp.173-186 (In Russian).</mixed-citation></citation-alternatives></ref><ref id="cit42"><label>42</label><citation-alternatives><mixed-citation xml:lang="ru">Nadachowski A. (1985). Biharian voles (Arvicolidae, Rodentia, Mammalia) from Kozi Grzbiet (Central Poland). Acta zool. Cracov., 29(2), pp. 13-28.</mixed-citation><mixed-citation xml:lang="en">Nadachowski A. (1985). Biharian voles (Arvicolidae, Rodentia, Mammalia) from Kozi Grzbiet (Central Poland). Acta zool. Cracov., 29(2), pp. 13-28.</mixed-citation></citation-alternatives></ref><ref id="cit43"><label>43</label><citation-alternatives><mixed-citation xml:lang="ru">Nadachowski, A. (1990). Review of fossil Rodentia from Poland (Mammalia). Senckenbergiana Biologica, 70(4/6), pp. 229-250.</mixed-citation><mixed-citation xml:lang="en">Nadachowski, A. (1990). Review of fossil Rodentia from Poland (Mammalia). Senckenbergiana Biologica, 70(4/6), pp. 229-250.</mixed-citation></citation-alternatives></ref><ref id="cit44"><label>44</label><citation-alternatives><mixed-citation xml:lang="ru">Nadachowski A., Žarski M., Urbanowski M., Wojtal P., Miekina B., Lipecki G., Ochman K., Krawczyk M., Jakubowski G. and Tomek T. (2009). Late Pleistocene Environment of the Czêstochowa Upland (Poland) reconstructed on the basis of faunistic evidence from archaeological cave sites. Kraków: Institute of Systematic and Evolution of Animals, Polish Academy of Sciences.</mixed-citation><mixed-citation xml:lang="en">Nadachowski A., Žarski M., Urbanowski M., Wojtal P., Miekina B., Lipecki G., Ochman K., Krawczyk M., Jakubowski G. and Tomek T. (2009). Late Pleistocene Environment of the Czêstochowa Upland (Poland) reconstructed on the basis of faunistic evidence from archaeological cave sites. Kraków: Institute of Systematic and Evolution of Animals, Polish Academy of Sciences.</mixed-citation></citation-alternatives></ref><ref id="cit45"><label>45</label><citation-alternatives><mixed-citation xml:lang="ru">Parfitt S. (1998). The interglacial mammalian fauna from Barnham. In: N. Ashton, S.G. Lewis and S. Parfitt, eds., Excavations at the Lower Palaeolithic Site at East Farm, Barnham: 1989-94. London: British Museum Occasional Paper, 125, pp. 111-147.</mixed-citation><mixed-citation xml:lang="en">Parfitt S. (1998). The interglacial mammalian fauna from Barnham. In: N. Ashton, S.G. Lewis and S. Parfitt, eds., Excavations at the Lower Palaeolithic Site at East Farm, Barnham: 1989-94. London: British Museum Occasional Paper, 125, pp. 111-147.</mixed-citation></citation-alternatives></ref><ref id="cit46"><label>46</label><citation-alternatives><mixed-citation xml:lang="ru">Pitts M. and Roberts M. (1997). Fairweather Eden: Life in Britain half a million years ago as revealed by the excavations at Boxgrove. London: http://en.wikipedia.org/wiki/Random_ House ISBN 9780712676861.</mixed-citation><mixed-citation xml:lang="en">Pitts M. and Roberts M. (1997). Fairweather Eden: Life in Britain half a million years ago as revealed by the excavations at Boxgrove. London: http://en.wikipedia.org/wiki/Random_ House ISBN 9780712676861.</mixed-citation></citation-alternatives></ref><ref id="cit47"><label>47</label><citation-alternatives><mixed-citation xml:lang="ru">Rekovets L.I. (1994). Anthropogene small mammals from the southern part of Eastern Europe. Kiev: Naukova Dumka Press (In Russian).</mixed-citation><mixed-citation xml:lang="en">Rekovets L.I. (1994). Anthropogene small mammals from the southern part of Eastern Europe. Kiev: Naukova Dumka Press (In Russian).</mixed-citation></citation-alternatives></ref><ref id="cit48"><label>48</label><citation-alternatives><mixed-citation xml:lang="ru">Schreve D.C. (2004a). The mammalian fauna of Barnfield Pit, Swanscombe, Kent. The Quaternary mammals of Southern and Eastern England. Field Guide. EuroMam UK, pp. 29 -48.</mixed-citation><mixed-citation xml:lang="en">Schreve D.C. (2004a). The mammalian fauna of Barnfield Pit, Swanscombe, Kent. The Quaternary mammals of Southern and Eastern England. Field Guide. EuroMam UK, pp. 29 -48.</mixed-citation></citation-alternatives></ref><ref id="cit49"><label>49</label><citation-alternatives><mixed-citation xml:lang="ru">Schreve D.C. (2004b). Late Middle Pleistocene (MIS 9) river Thames terrace deposits at Purfleet, Essex. The Quaternary mammals of Southern &amp; Eastern England. Field Guide. EuroMam UK, pp. 49 -67.</mixed-citation><mixed-citation xml:lang="en">Schreve D.C. (2004b). Late Middle Pleistocene (MIS 9) river Thames terrace deposits at Purfleet, Essex. The Quaternary mammals of Southern &amp; Eastern England. Field Guide. EuroMam UK, pp. 49 -67.</mixed-citation></citation-alternatives></ref><ref id="cit50"><label>50</label><citation-alternatives><mixed-citation xml:lang="ru">Schreve D.C. (2004c). The mammalian fauna of the penultimate (MIS 7) interglacial in the lower Themes valley. The Quaternary mammals of Southern and Eastern England. Field Guide. EuroMam UK, pp. 69 – 79.</mixed-citation><mixed-citation xml:lang="en">Schreve D.C. (2004c). The mammalian fauna of the penultimate (MIS 7) interglacial in the lower Themes valley. The Quaternary mammals of Southern and Eastern England. Field Guide. EuroMam UK, pp. 69 – 79.</mixed-citation></citation-alternatives></ref><ref id="cit51"><label>51</label><citation-alternatives><mixed-citation xml:lang="ru">Shik S.M. (2014). Neopleistocene of central European Russia (the new ideas on stratigraphy and palaeogeography). Stratigraphy. Geol. Correlation, 22 (2), pp. 108-120 (In Russian).</mixed-citation><mixed-citation xml:lang="en">Shik S.M. (2014). Neopleistocene of central European Russia (the new ideas on stratigraphy and palaeogeography). Stratigraphy. Geol. Correlation, 22 (2), pp. 108-120 (In Russian).</mixed-citation></citation-alternatives></ref><ref id="cit52"><label>52</label><citation-alternatives><mixed-citation xml:lang="ru">Siori M.S. and Sala B. (2007). The mammal fauna from the late Early Biharian site of Castagnone (Northern Monferrato, Piedmant, NW Italy). Geobios, 40, pp. 207-217.</mixed-citation><mixed-citation xml:lang="en">Siori M.S. and Sala B. (2007). The mammal fauna from the late Early Biharian site of Castagnone (Northern Monferrato, Piedmant, NW Italy). Geobios, 40, pp. 207-217.</mixed-citation></citation-alternatives></ref><ref id="cit53"><label>53</label><citation-alternatives><mixed-citation xml:lang="ru">Socha P. (2014). Rodent palaeofaunas from Biśik Cave (Krakow-Częstochowa Upland, Poland): palaeoecological, palaeoclimatic and biostrtigraphic reconstruction. Quaternary International. 326-327, pp. 64-81.</mixed-citation><mixed-citation xml:lang="en">Socha P. (2014). Rodent palaeofaunas from Biśik Cave (Krakow-Częstochowa Upland, Poland): palaeoecological, palaeoclimatic and biostrtigraphic reconstruction. Quaternary International. 326-327, pp. 64-81.</mixed-citation></citation-alternatives></ref><ref id="cit54"><label>54</label><citation-alternatives><mixed-citation xml:lang="ru">Stuart A.J. and Lister A.M. (2001). The mammalian faunas of Pakefield/Kessingland and Corton, Suffolk, UK: evidence for a new temperate episode in the British early Middle Pleistocene. Quaternary Science Reviews, 20, pp. 1677–1692.</mixed-citation><mixed-citation xml:lang="en">Stuart A.J. and Lister A.M. (2001). The mammalian faunas of Pakefield/Kessingland and Corton, Suffolk, UK: evidence for a new temperate episode in the British early Middle Pleistocene. Quaternary Science Reviews, 20, pp. 1677–1692.</mixed-citation></citation-alternatives></ref><ref id="cit55"><label>55</label><citation-alternatives><mixed-citation xml:lang="ru">Van Kolfschoten T. (2014). The Palaeolithic locality Schöningen (Germany): A review of the mammalian record. Quaternary International. 326-327, pp. 469-480.</mixed-citation><mixed-citation xml:lang="en">Van Kolfschoten T. (2014). The Palaeolithic locality Schöningen (Germany): A review of the mammalian record. Quaternary International. 326-327, pp. 469-480.</mixed-citation></citation-alternatives></ref><ref id="cit56"><label>56</label><citation-alternatives><mixed-citation xml:lang="ru">Van Kolfschoten T. (1990). Review of the Pleistocene arvicolid faunas from the Netherlands. Prague: Int. Symp. Evol. Phyl. Biostr. Arvicolids, pp. 255-274.</mixed-citation><mixed-citation xml:lang="en">Van Kolfschoten T. (1990). Review of the Pleistocene arvicolid faunas from the Netherlands. Prague: Int. Symp. Evol. Phyl. Biostr. Arvicolids, pp. 255-274.</mixed-citation></citation-alternatives></ref><ref id="cit57"><label>57</label><citation-alternatives><mixed-citation xml:lang="ru">Van Kolfschoten T. and Turner E. (1996). Early Middle Pleistocene Faunas from Kärlich and Miesenheim I and their biostratigraphic implication. In: Turner C., ed., The early Middle Pleistocene in Europe. Balkema, Rotterdam, pp. 227–253.</mixed-citation><mixed-citation xml:lang="en">Van Kolfschoten T. and Turner E. (1996). Early Middle Pleistocene Faunas from Kärlich and Miesenheim I and their biostratigraphic implication. In: Turner C., ed., The early Middle Pleistocene in Europe. Balkema, Rotterdam, pp. 227–253.</mixed-citation></citation-alternatives></ref><ref id="cit58"><label>58</label><citation-alternatives><mixed-citation xml:lang="ru">Van der Meulen A.J. (1973). Middle Pleistocene smaller mammals from the Monte Peglia (Oriento, Italy) with special reference to the phylogeny of Microtus (Arvicolidae, Rodentia). Quaternaria, 17, pp. 1-144.</mixed-citation><mixed-citation xml:lang="en">Van der Meulen A.J. (1973). Middle Pleistocene smaller mammals from the Monte Peglia (Oriento, Italy) with special reference to the phylogeny of Microtus (Arvicolidae, Rodentia). Quaternaria, 17, pp. 1-144.</mixed-citation></citation-alternatives></ref><ref id="cit59"><label>59</label><citation-alternatives><mixed-citation xml:lang="ru">Velichko A.A., Markova A.K., Morozova T.D., Nechaev V.P., Svetlitckaya T.V., Tcatckin A.I. and Chichagova O.A. (1992). The geochronology of loess-soil formation S-W of the Russian Plain by new data. Geochronology of the Quaternary. Moscow: Nauka Press, pp. 28-33 (In Russian).</mixed-citation><mixed-citation xml:lang="en">Velichko A.A., Markova A.K., Morozova T.D., Nechaev V.P., Svetlitckaya T.V., Tcatckin A.I. and Chichagova O.A. (1992). The geochronology of loess-soil formation S-W of the Russian Plain by new data. Geochronology of the Quaternary. Moscow: Nauka Press, pp. 28-33 (In Russian).</mixed-citation></citation-alternatives></ref><ref id="cit60"><label>60</label><citation-alternatives><mixed-citation xml:lang="ru">Vislobokova I. and Tesakov A. (2013). Early and Middle Pleistocene of Northern Eurasia. In: Elias S.A., ed., The Encyclopedia of Quaternary Science, vol. 4. Amsterdam: Elsevier, pp. 605-614.</mixed-citation><mixed-citation xml:lang="en">Vislobokova I. and Tesakov A. (2013). Early and Middle Pleistocene of Northern Eurasia. In: Elias S.A., ed., The Encyclopedia of Quaternary Science, vol. 4. Amsterdam: Elsevier, pp. 605-614.</mixed-citation></citation-alternatives></ref><ref id="cit61"><label>61</label><citation-alternatives><mixed-citation xml:lang="ru">Von Koenigswald W. and Heinrich, W.-D. (1999). Mittelpleistozäne Säugetierfaunen aus Mitteleuropa – der Versuch einer biostratigraphischen Zuordnung. Darmstädter Beiträge zur Naturgeschichte, 9, pp. 53-112.</mixed-citation><mixed-citation xml:lang="en">Von Koenigswald W. and Heinrich, W.-D. (1999). Mittelpleistozäne Säugetierfaunen aus Mitteleuropa – der Versuch einer biostratigraphischen Zuordnung. Darmstädter Beiträge zur Naturgeschichte, 9, pp. 53-112.</mixed-citation></citation-alternatives></ref><ref id="cit62"><label>62</label><citation-alternatives><mixed-citation xml:lang="ru">Von Koenigswald W. and van Kolfschoten, T. (1996). The Mimomys – Arvicola boundary and the enamel thickness quotient (SDQ) of Arvicola as stratigraphic markers in the Middle Pleistocene. In. Turner C., ed., The early Middle Pleistocene in Europe. Balkema, Rotterdam, pp. 211-226.</mixed-citation><mixed-citation xml:lang="en">Von Koenigswald W. and van Kolfschoten, T. (1996). The Mimomys – Arvicola boundary and the enamel thickness quotient (SDQ) of Arvicola as stratigraphic markers in the Middle Pleistocene. In. Turner C., ed., The early Middle Pleistocene in Europe. Balkema, Rotterdam, pp. 211-226.</mixed-citation></citation-alternatives></ref><ref id="cit63"><label>63</label><citation-alternatives><mixed-citation xml:lang="ru">Zastrozhnov A., Danukalova G. and Murray A. (2017). New data on the age of the Neopleistocene deposits of the lower Volga sites according to the OSL method results. Fundamental problems of Quaternary: the results of investigation and the main directions in the future. Moscow: GEOS, pp. 139-140.</mixed-citation><mixed-citation xml:lang="en">Zastrozhnov A., Danukalova G. and Murray A. (2017). New data on the age of the Neopleistocene deposits of the lower Volga sites according to the OSL method results. Fundamental problems of Quaternary: the results of investigation and the main directions in the future. Moscow: GEOS, pp. 139-140.</mixed-citation></citation-alternatives></ref></ref-list><fn-group><fn fn-type="conflict"><p>The authors declare that there are no conflicts of interest present.</p></fn></fn-group></back></article>
